The title “ecologies of change” might seem paradoxical to some; to others even tautological. Is ecology synonymous with the lasting and unchanging and thus in urgent need of preservation, to be defended against human interventions and technological change? Or is change inherent to ecologies and thus an ecological mode of thinking puts forward a dynamic, procedural and open way to conceptualise the world? Homeostasis, or the self-regulation of nature presupposes a concept of nature as separate from culture. Thus nature always already is a discursive construct, onto which ideals of regulation and (self) control are projected. There is no easy or singular answer to the question of what media ecology is. The contributions in this issue of spheres touch upon this plurality and are concerned with the concept of (media) ecologies in diverse ways.